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26/01/2026 Island restoration

Darwin and the land iguanas of Santiago

Charles Darwin was one of the last people to describe land iguanas living on Santiago and was not exactly polite about these reptiles. But it wasn’t long until Darwin's inquisitive nature took hold...

Photograph of Henry Nicholls

Henry Nicholls

Henry Nicholls is a GCT Ambassador and editor of 'Galapagos Matters' magazine. He works as a secondary school teacher but has also spent many years as a freelance science journalist specialising in evolutionary biology, the environment, conservation and history of science. He has written two books about Galapagos, 'Lonesome George: The Life and Loves of a Conservation Icon' and 'The Galapagos: A Natural History'.

When Charles Darwin met the land iguanas living on Santiago, he wasn’t very impressed. “From their low facial angle they have a singularly stupid appearance,” he wrote. “In their movements, they are lazy and half torpid.” There were so many of them nesting on the lower slopes of Santiago that Darwin and his Beagle shipmates “could not for some time find a spot free from their burrows” on which to pitch a tent. But once settled, Darwin turned his inquisitive mind to finding out everything he could about their biology. He cut open a few specimens, finding their stomachs “full of vegetable fibres and leaves of different trees” and, observing that they were “very fond of cactus”, he began to feed them with slices of Optuntia. “When a piece is thrown towards them, each will try to seize & carry it away as dogs do with a bone,” he wrote.

Galapagos land iguana on North Seymour
Galapagos land iguana © Kenneth Arrison

In their movements they are lazy and half torpid.

Charles Darwin

Santiago’s land iguanas, like almost every other animal in Galapagos, did not appear to be fearful. In a confrontation, either with another iguana or Darwin himself, they would curl their tails, raise themselves on their forelimbs, and bob their heads up and down quickly, which “gives them rather a fierce aspect.” When Darwin poked one with a stick, he found it would “bite it severely” and that two animals “placed on the ground close together” would fight “till blood is drawn.” When the Beagle left the Galapagos a few weeks later, it sailed away with one land iguana specimen pickled in “Spirits of Wine”.

Galapagos land iguana yawning
Galapagos land iguana © Jason Lim

Darwin in Galapagos

Discover more about Darwin’s historic visit to the Galapagos Islands.

Find out more

This account and a few others made by naturalists before and just after Darwin are the only records we have of land iguanas living on Santiago. Over the next few decades, pressures exerted by increasing human footfall – including further disruption of nesting grounds, the harvesting of iguanas and their eggs for food and the introduction of pigs and goats to Santiago –led to the extinction of the island’s land iguanas. Yet today, in 2026, more than 150 years after their disappearance from this island, the land iguanas are making a comeback.

Santiago land iguana project, Galapagos
Santiago land iguana project © Johannes Ramírez
Land iguana on North Seymour island, Galapagos

Reintroducing land iguanas to Santiago

Discover the groundbreaking work to return of the Galapagos land iguana to Santiago.

Find out more

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